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The Kuparuk River, an arctic stream ecosystem on the North Slope of Alaska.The Kuparuk River flows to the Arctic Ocean from the foothills of the Brooks Range, seen to the south in the background. The Kuparuk River was experimentally enriched with phosphorus every summer from 1983 to 1998 as a part of the Toolik Lake Long-Term Ecological Research (LTER) project. The purpose of the experiment was to evaluate the long-term effects of eutrophication on an arctic tundra stream. During the 16 years of phosphorus additions, a dramatic change was observed in the community structure and at all trophic levels. This photograph originally appeared on the cover of Ecology (85:4) in April of 2004.
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